About the Authors

Luis Arnes

Contributed equally to this work with: Luis Arnes, Jonathon T. Hill

Affiliation Department of Genetics and Development, Columbia University, New York, New York, United States of America

Jonathon T. Hill

Contributed equally to this work with: Luis Arnes, Jonathon T. Hill

Affiliation Department of Genetics and Development, Columbia University, New York, New York, United States of America

Stefanie Gross

Affiliation Department of Genetics and Development, Columbia University, New York, New York, United States of America

Mark A. Magnuson

Affiliation Department of Molecular Physiology and Biophysics and Center for Stem Cell Biology, Vanderbilt University School of Medicine, Nashville, Tennessee, United States of America

Lori Sussel

lgs2@columbia.edu

Affiliation Department of Genetics and Development, Columbia University, New York, New York, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: LS LA JH. Performed the experiments: LS LA JH SG. Contributed reagents/materials/analysis tools: MM. Wrote the paper: LS LA JH.