About the Authors

Giichi Takaesu

takaesug1@z7.keio.jp (GT); Jun_Tsuji@ncsu.edu (JN-T)

Affiliation Center for Integrated Medical Research, Keio University School of Medicine, Tokyo, Japan

Maiko Inagaki

Current address: Research Institute for Radiation Biology and Medicine, Hiroshima University, Hiroshima, Japan

Affiliation Department of Environmental and Molecular Toxicology, North Carolina State University, Raleigh, North Carolina, United States of America

Keiyo Takubo

Affiliation Department of Cell Differentiation, The Sakaguchi Laboratory of Developmental Biology, Keio University School of Medicine, Tokyo, Japan

Yuji Mishina

Affiliation Department of Biologic and Materials Sciences, School of Dentistry, University of Michigan, Ann Arbor, Michigan, United States of America

Paul R. Hess

Affiliation Department of Clinical Sciences, College of Veterinary Medicine, North Carolina State University, Raleigh, North Carolina, United States of America

Gregg A. Dean

Affiliations Department of Clinical Sciences, College of Veterinary Medicine, North Carolina State University, Raleigh, North Carolina, United States of America, Center for Comparative Medicine and Translational Research, College of Veterinary Medicine, North Carolina State University, Raleigh, North Carolina, United States of America

Akihiko Yoshimura

Affiliation Department of Microbiology and Immunology, Keio University School of Medicine, Tokyo, Japan

Kunihiro Matsumoto

Affiliation Department of Molecular Biology, Graduate School of Science, Nagoya University, Nagoya, Japan

Toshio Suda

Affiliation Department of Cell Differentiation, The Sakaguchi Laboratory of Developmental Biology, Keio University School of Medicine, Tokyo, Japan

Jun Ninomiya-Tsuji

takaesug1@z7.keio.jp (GT); Jun_Tsuji@ncsu.edu (JN-T)

Affiliation Department of Environmental and Molecular Toxicology, North Carolina State University, Raleigh, North Carolina, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: GT YM PH GD AY KM TS JN-T. Performed the experiments: GT MI KT. Analyzed the data: GT JN-T. Wrote the paper: GT JN-T.