About the Authors

Kei Watanabe

Contributed equally to this work with: Kei Watanabe, Kenta Wada

Affiliations Mammalian Genetics Project, Tokyo Metropolitan Institute of Medical Science, Tokyo, Japan, Graduate School of Life and Environmental Sciences, University of Tsukuba, Tsukuba, Japan

Kenta Wada

Contributed equally to this work with: Kei Watanabe, Kenta Wada

Affiliations Mammalian Genetics Project, Tokyo Metropolitan Institute of Medical Science, Tokyo, Japan, Department of Bioproduction, Tokyo University of Agriculture, Abashiri, Japan

Tomoko Ohashi

Current address: Department of Medical Genome Sciences, Research Institute for Frontier Medicine, Sapporo Medical University, School of Medicine, Sapporo, Japan

Affiliation Department of Bioproduction, Tokyo University of Agriculture, Abashiri, Japan

Saki Okubo

Affiliation Department of Bioproduction, Tokyo University of Agriculture, Abashiri, Japan

Kensuke Takekuma

Affiliation Department of Bioproduction, Tokyo University of Agriculture, Abashiri, Japan

Ryoichi Hashizume

Affiliation Department of Bioproduction, Tokyo University of Agriculture, Abashiri, Japan

Jun-Ichi Hayashi

Affiliation Graduate School of Life and Environmental Sciences, University of Tsukuba, Tsukuba, Japan

Tadao Serikawa

Affiliation Institute of Laboratory Animals, Graduate School of Medicine, Kyoto University, Kyoto, Japan

Takashi Kuramoto

Affiliation Institute of Laboratory Animals, Graduate School of Medicine, Kyoto University, Kyoto, Japan

Yoshiaki Kikkawa

kikkawa-ys@igakuken.or.jp

Affiliation Mammalian Genetics Project, Tokyo Metropolitan Institute of Medical Science, Tokyo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: K. Watanabe K. Wada J-IH TS TK YK. Performed the experiments: K. Watanabe K. Wada TO SO KT RH TK YK. Analyzed the data: K. Watanabe K. Wada TO TK YK. Contributed reagents/materials/analysis tools: TS TK. Wrote the paper: K. Watanabe K. Wada J-IH TS TK YK.