About the Authors

Annette Roug

Affiliation Wildlife Health Center, School of Veterinary Medicine, University of California Davis, Davis, California, United States of America

Pamela Swift

Affiliation California Department of Fish and Game, Rancho Cordova, California, United States of America

Steven Torres

Affiliation California Department of Fish and Game, Rancho Cordova, California, United States of America

Karen Jones

Affiliation California Department of Fish and Game, Rancho Cordova, California, United States of America

Christine K. Johnson

ckjohnson@ucdavis.edu

Affiliation Wildlife Health Center, School of Veterinary Medicine, University of California Davis, Davis, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: AR PS ST KJ CKJ. Performed the experiments: AR PS ST KJ. Analyzed the data: AR CKJ. Contributed reagents/materials/analysis tools: PS ST KJ CKJ. Wrote the paper: AR PS CKJ.