About the Authors

Xiong-jian Luo

Contributed equally to this work with: Xiong-jian Luo, Ming Li

Affiliations State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, Yunnan, China, University of Rochester Flaum Eye Institute, University of Rochester, Rochester, New York, United States of America

Ming Li

Contributed equally to this work with: Xiong-jian Luo, Ming Li

Affiliation State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, Yunnan, China

Liang Huang

Affiliations Gannan Medical University, Ganzhou, Jiangxi, China, University of Rochester Flaum Eye Institute, University of Rochester, Rochester, New York, United States of America

Kwangsik Nho

Affiliations Center for Neuroimaging, Department of Radiology and Imaging Sciences, Indiana University School of Medicine, Indianapolis, Indiana, United States of America, Center for Computational Biology and Bioinformatics, Indiana University School of Medicine, Indianapolis, Indiana, United States of America, Regenstrief Institute, Indianapolis, Indiana, United States of America, Department of Medical and Molecular Genetics, Indiana University School of Medicine, Indianapolis, Indiana, United States of America

Min Deng

Affiliation University of Rochester Flaum Eye Institute, University of Rochester, Rochester, New York, United States of America

Qiang Chen

Affiliation Genes, Cognition and Psychosis Program, National Institute of Mental Health, the National Institutes of Health, Bethesda, Maryland, United States of America

Daniel R. Weinberger

Affiliation Genes, Cognition and Psychosis Program, National Institute of Mental Health, the National Institutes of Health, Bethesda, Maryland, United States of America

Alejandro Arias Vasquez

Affiliations Department of Genetics, Radboud University Nijmegen Medical Centre, Nijmegen, The Netherlands, Department of Psychiatry, Donders Institute for Brain, Cognition and Behaviour, Radboud University Nijmegen Medical Centre, Nijmegen, The Netherlands

Mark Rijpkema

Affiliation Donders Institute for Brain, Cognition and Behaviour, Donders Centre for Cognitive Neuroimaging, Radboud University Nijmegen, Nijmegen, The Netherlands

Venkata S. Mattay

Affiliation Genes, Cognition and Psychosis Program, National Institute of Mental Health, the National Institutes of Health, Bethesda, Maryland, United States of America

Andrew J. Saykin

Affiliations Center for Neuroimaging, Department of Radiology and Imaging Sciences, Indiana University School of Medicine, Indianapolis, Indiana, United States of America, Regenstrief Institute, Indianapolis, Indiana, United States of America, Department of Medical and Molecular Genetics, Indiana University School of Medicine, Indianapolis, Indiana, United States of America

Li Shen

Affiliations Center for Neuroimaging, Department of Radiology and Imaging Sciences, Indiana University School of Medicine, Indianapolis, Indiana, United States of America, Regenstrief Institute, Indianapolis, Indiana, United States of America, Department of Medical and Molecular Genetics, Indiana University School of Medicine, Indianapolis, Indiana, United States of America

Guillén Fernández

Affiliations Donders Institute for Brain, Cognition and Behaviour, Donders Centre for Cognitive Neuroimaging, Radboud University Nijmegen, Nijmegen, The Netherlands, Department of Cognitive Neuroscience, Donders Institute for Brain, Cognition and Behaviour, Radboud University Nijmegen Medical Centre, Nijmegen, The Netherlands

Barbara Franke

Affiliations Department of Genetics, Radboud University Nijmegen Medical Centre, Nijmegen, The Netherlands, Department of Psychiatry, Donders Institute for Brain, Cognition and Behaviour, Radboud University Nijmegen Medical Centre, Nijmegen, The Netherlands

Jing-chun Chen

Affiliation Virginia Institute for Psychiatric and Behavioral Genetics, and Department of Psychiatry, Virginia Commonwealth University, Richmond, Virginia, United States of America

Xiang-ning Chen

Affiliation Virginia Institute for Psychiatric and Behavioral Genetics, and Department of Psychiatry, Virginia Commonwealth University, Richmond, Virginia, United States of America

Jin-kai Wang

Affiliation State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, Yunnan, China

Xiao Xiao

Affiliation State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, Yunnan, China

Xue-bin Qi

Affiliation State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, Yunnan, China

Kun Xiang

Affiliation State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, Yunnan, China

Ying-Mei Peng

Affiliation State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, Yunnan, China

Xiang-yu Cao

Affiliation State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, Yunnan, China

Yi Li

Affiliation Biological Resources and Environmental Science College, Qujing Normal University, Qujing, Yunnan, China

Xiao-dong Shi

Affiliation Biological Resources and Environmental Science College, Qujing Normal University, Qujing, Yunnan, China

for the Alzheimer’s Disease Neuroimaging Initiative
Lin Gan

sub@mail.kiz.ac.cn (BS); Lin_Gan@urmc.rochester.edu (LG)

Affiliations University of Rochester Flaum Eye Institute, University of Rochester, Rochester, New York, United States of America, College of Life and Environmental Sciences, Hangzhou Normal University, Hangzhou, Zhejiang, China

Bing Su

sub@mail.kiz.ac.cn (BS); Lin_Gan@urmc.rochester.edu (LG)

Affiliation State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, Yunnan, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: XJL ML LG BS. Performed the experiments: XJL ML LH MD. Analyzed the data: XJL ML KN QC DRW AAV MR VSM AJS LS GF BF JCC XNC. Contributed reagents/materials/analysis tools: XX JKW XBQ KX YMP XYC YL XDS. Wrote the paper: XJL ML LG BS.