About the Authors

Helen C. Johnson

helen.johnson@lshtm.ac.uk

Affiliation Department of Infectious Disease Epidemiology/Centre for the Mathematical Modelling of Infectious Diseases, London School of Hygiene and Tropical Medicine, London, United Kingdom

K. Miriam Elfström

Affiliation Department of Medical Epidemiology and Biostatistics, Karolinska Institutet, Stockholm, Sweden

W. John Edmunds

Affiliation Department of Infectious Disease Epidemiology/Centre for the Mathematical Modelling of Infectious Diseases, London School of Hygiene and Tropical Medicine, London, United Kingdom

Competing Interests

WJE’s partner works for GSK. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: HCJ WJE. Performed the experiments: HCJ. Analyzed the data: HCJ KME. Wrote the paper: HCJ.