About the Authors

Naoko Goto-Inoue

Affiliation Department of Cell Biology and Anatomy, Hamamatsu University School of Medicine, 1-20-1 Handayama, Higashi-ku, Hamamatsu, Shizuoka, Japan

Takahiro Hayasaka

Affiliation Department of Cell Biology and Anatomy, Hamamatsu University School of Medicine, 1-20-1 Handayama, Higashi-ku, Hamamatsu, Shizuoka, Japan

Nobuhiro Zaima

Affiliation Department of Applied Biological Chemistry, Kinki University, Nara, Nara, Japan

Kimiko Nakajima

Affiliation Department of Dermatology, Kochi Medical School, Kochi University, Kohasu, Okocho, Nankoku, Nankoku, Japan

Walter M. Holleran

Affiliation Department of Dermatology, School of Medicine, University of California San Francisco, Department of Veterans Affairs Medical Center, and Northern California Institute for Research and Education, San Francisco, California, United States of America

Shigetoshi Sano

Affiliation Department of Dermatology, Kochi Medical School, Kochi University, Kohasu, Okocho, Nankoku, Nankoku, Japan

Yoshikazu Uchida

Contributed equally to this work with: Yoshikazu Uchida, Mitsutoshi Setou

uchiday@derm.ucsf.edu (YU); setou@hama-med.ac.jp (MS)

Affiliation Department of Dermatology, School of Medicine, University of California San Francisco, Department of Veterans Affairs Medical Center, and Northern California Institute for Research and Education, San Francisco, California, United States of America

Mitsutoshi Setou

Contributed equally to this work with: Yoshikazu Uchida, Mitsutoshi Setou

uchiday@derm.ucsf.edu (YU); setou@hama-med.ac.jp (MS)

Affiliation Department of Cell Biology and Anatomy, Hamamatsu University School of Medicine, 1-20-1 Handayama, Higashi-ku, Hamamatsu, Shizuoka, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: NG-I MS YU. Performed the experiments: NG-I. Analyzed the data: NG-I TH. Contributed reagents/materials/analysis tools: NZ KN WM-H SS. Wrote the paper: NG-I MS YU.

. Analyzed the data: NG-I TH. Contributed reagents/materials/analysis tools: NZ KN WM-H SS. Wrote the paper: NG-I MS YU.