About the Authors

Gilles Crevel

Affiliation Department Basic Medical Sciences, St. Georges University London, Cranmer Terrace, London, United Kingdom

Nicole Vo

Affiliation Department Basic Medical Sciences, St. Georges University London, Cranmer Terrace, London, United Kingdom

Isabelle Crevel

Affiliation Department Basic Medical Sciences, St. Georges University London, Cranmer Terrace, London, United Kingdom

Sana Hamid

Affiliation Department Basic Medical Sciences, St. Georges University London, Cranmer Terrace, London, United Kingdom

Lily Hoa

Affiliation Department Basic Medical Sciences, St. Georges University London, Cranmer Terrace, London, United Kingdom

Seiji Miyata

Affiliations Department Basic Medical Sciences, St. Georges University London, Cranmer Terrace, London, United Kingdom, Department of Applied Biology, Kyoto Institute of Technology, Kyoto, Japan

Sue Cotterill

s.cotterill@sgul.ac.uk

Affiliation Department Basic Medical Sciences, St. Georges University London, Cranmer Terrace, London, United Kingdom

Competing Interests

Co-author Sue Cotterill is a PLOS ONE Editorial Board member. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: SC GC. Performed the experiments: GC IC NV LH SH SC. Analyzed the data: SC GC IC. Contributed reagents/materials/analysis tools: SM. Wrote the paper: SC GC.