About the Authors

Tabitha A. Graves

Tabitha.Graves@colostate.edu

Affiliation School of Forestry, Northern Arizona University, Flagstaff, Arizona, United States of America

J. Andrew Royle

Affiliation United States Geological Survey, Patuxent Wildlife Research Center, Laurel, Maryland, United States of America

Katherine C. Kendall

Affiliation United States Geological Survey, Northern Rocky Mountain Science Center, Glacier Field Station, Glacier National Park, Montana, United States of America

Paul Beier

Affiliation School of Forestry, Northern Arizona University, Flagstaff, Arizona, United States of America

Jeffrey B. Stetz

Affiliation Montana Cooperative Wildlife Research Unit, University of Montana, Missoula, Montana, United States of America

Amy C. Macleod

Affiliation University of Montana Cooperative Ecosystem Studies Unit, Glacier Field Station, Glacier National Park, Montana, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TAG JAR KCK PB JBS. Performed the experiments: TAG KCK JBS ACM. Analyzed the data: TAG JAR KCK. Contributed reagents/materials/analysis tools: TAG JAR KCK. Wrote the paper: TAG JAR KCK PB JBS ACM.