About the Authors

Johann Louarn

Affiliations Université de Toulouse; UPS; UMR 5546, Laboratoire de Recherche en Sciences Végétales; BP 42617 Auzeville, F-31326 Castanet-Tolosan, France, CNRS; UMR 5546; BP 42617, F-31326 Castanet-Tolosan, France

Francis Carbonne

Affiliations Université de Toulouse; UPS; UMR 5546, Laboratoire de Recherche en Sciences Végétales; BP 42617 Auzeville, F-31326 Castanet-Tolosan, France, CNRS; UMR 5546; BP 42617, F-31326 Castanet-Tolosan, France

Philippe Delavault

Affiliation Laboratoire de Biologie et Pathologie Végétales, SFR 4207 QUASAV, LUNAM University, Nantes, France

Guillaume Bécard

Affiliations Université de Toulouse; UPS; UMR 5546, Laboratoire de Recherche en Sciences Végétales; BP 42617 Auzeville, F-31326 Castanet-Tolosan, France, CNRS; UMR 5546; BP 42617, F-31326 Castanet-Tolosan, France

Soizic Rochange

rochange@scsv.ups-tlse.fr

Affiliations Université de Toulouse; UPS; UMR 5546, Laboratoire de Recherche en Sciences Végétales; BP 42617 Auzeville, F-31326 Castanet-Tolosan, France, CNRS; UMR 5546; BP 42617, F-31326 Castanet-Tolosan, France

Competing Interests

This research was funded by ‘Soltis’ and ‘Syngenta’. There are no patents, products in development or marketed products to declare. This does not alter the authors’ adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: JL PD GB SR. Performed the experiments: JL FC. Analyzed the data: JL FC PD GB SR. Contributed reagents/materials/analysis tools: PD. Wrote the paper: JL PD GB SR.