About the Authors

Stephanie Georgina Dakin

sdakin@rvc.ac.uk

Affiliation Department of Veterinary Clinical Sciences, Royal Veterinary College, University of London, Hatfield, United Kingdom

Jayesh Dudhia

Affiliation Department of Veterinary Clinical Sciences, Royal Veterinary College, University of London, Hatfield, United Kingdom

Natalie Jayne Werling

Affiliation Department of Biotherapeutics, National Institute for Biological Standards and Control, South Mimms, United Kingdom

Dirk Werling

Affiliation Department of Pathology and Infectious Diseases, Royal Veterinary College, University of London, Hatfield, United Kingdom

Dilkush Robert Ephrem Abayasekara

Affiliation Department of Veterinary Basic Sciences, Royal Veterinary College, University of London, London, United Kingdom

Roger Kenneth Whealands Smith

Affiliation Department of Veterinary Clinical Sciences, Royal Veterinary College, University of London, Hatfield, United Kingdom

Competing Interests

The authors declare the study was in part funded by Ceva (France). There are no other relevant declarations relating to employment, consultancy, patents, products in development or marketed products. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: SGD JD DREA RKWS. Performed the experiments: SGD. Analyzed the data: SGD JD NJW RKWS. Contributed reagents/materials/analysis tools: SGD JD RKWS. Wrote the paper: SGD JD NJW DW DREA RKWS.