About the Authors

Takashi Chaen

Contributed equally to this work with: Takashi Chaen, Toshihiro Konno

Affiliation Laboratory of Animal Breeding, Department of Veterinary Medical Sciences, The University of Tokyo, Bunkyo-ku, Tokyo, Japan

Toshihiro Konno

Contributed equally to this work with: Takashi Chaen, Toshihiro Konno

konnot@gmail.com

Affiliation Laboratory of Animal Breeding, Department of Veterinary Medical Sciences, The University of Tokyo, Bunkyo-ku, Tokyo, Japan

Mahiro Egashira

Affiliation Laboratory of Animal Breeding, Department of Veterinary Medical Sciences, The University of Tokyo, Bunkyo-ku, Tokyo, Japan

Rulan Bai

Affiliation Laboratory of Animal Breeding, Department of Veterinary Medical Sciences, The University of Tokyo, Bunkyo-ku, Tokyo, Japan

Nana Nomura

Affiliation Laboratory of Animal Breeding, Department of Veterinary Medical Sciences, The University of Tokyo, Bunkyo-ku, Tokyo, Japan

Shintaro Nomura

Affiliation Department of Animal Bioscience, Nagahama Institute of Bio-Science and Technology, Nagahama, Shiga, Japan

Yasushi Hirota

Affiliation Department of Obstetrics and Gynecology, Faculty of Medicine, The University of Tokyo, Bunkyo-ku, Tokyo, Japan

Toshihiro Sakurai

Affiliation Laboratory of Animal Breeding, Department of Veterinary Medical Sciences, The University of Tokyo, Bunkyo-ku, Tokyo, Japan

Kazuhiko Imakawa

Affiliation Laboratory of Animal Breeding, Department of Veterinary Medical Sciences, The University of Tokyo, Bunkyo-ku, Tokyo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TK TC. Performed the experiments: TC TK ME RB NN SN YH TS. Analyzed the data: TC TK ME. Contributed reagents/materials/analysis tools: KI. Wrote the paper: TK TC.