About the Authors

Richard D. Antrobus

Contributed equally to this work with: Richard D. Antrobus, Patrick J. Lillie, Tamara K. Berthoud

Affiliation The Jenner Institute, University of Oxford, Oxford, United Kingdom

Patrick J. Lillie

Contributed equally to this work with: Richard D. Antrobus, Patrick J. Lillie, Tamara K. Berthoud

Affiliation The Jenner Institute, University of Oxford, Oxford, United Kingdom

Tamara K. Berthoud

Contributed equally to this work with: Richard D. Antrobus, Patrick J. Lillie, Tamara K. Berthoud

Affiliation The Jenner Institute, University of Oxford, Oxford, United Kingdom

Alexandra J. Spencer

Affiliation The Jenner Institute, University of Oxford, Oxford, United Kingdom

James E. McLaren

Affiliation Institute of Infection and Immunity, Cardiff University School of Medicine, Cardiff, United Kingdom

Kristin Ladell

Affiliation Institute of Infection and Immunity, Cardiff University School of Medicine, Cardiff, United Kingdom

Teresa Lambe

Affiliation The Jenner Institute, University of Oxford, Oxford, United Kingdom

Anita Milicic

Affiliation The Jenner Institute, University of Oxford, Oxford, United Kingdom

David A. Price

Affiliation Institute of Infection and Immunity, Cardiff University School of Medicine, Cardiff, United Kingdom

Adrian V. S. Hill

Affiliation The Jenner Institute, University of Oxford, Oxford, United Kingdom

Sarah C. Gilbert

sarah.gilbert@ndm.ox.ac.uk

Affiliation The Jenner Institute, University of Oxford, Oxford, United Kingdom

Competing Interests

SCG and AVSH are named as inventors on patents relating to methods of vaccination, including influenza vaccines: Methods and reagents for vaccination which generate a CD8+ T cell response. 1997 06 07. United States Patent Application 20110159034. Compositions and methods 2007 10 05. United States Patent Application 20100285050. All other authors have declared that no competing interests exist. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: TL AM DP AVSH SCG. Performed the experiments: RA PL TKB AJS JM KL. Analyzed the data: RA TKB JM Kl DP. Wrote the paper: RA TKB DP SCG.