About the Authors

Karen H. Black

k.black@unsw.edu.au (KHB); aaron.b.camens@gmail.com (ABC)

Affiliation School of Biological, Earth and Environmental Sciences, University of New South Wales, Sydney, New South Wales, Australia

Aaron B. Camens

k.black@unsw.edu.au (KHB); aaron.b.camens@gmail.com (ABC)

Affiliation Department of Earth and Environmental Sciences, University of Adelaide, Adelaide, South Australia, Australia

Michael Archer

Affiliation School of Biological, Earth and Environmental Sciences, University of New South Wales, Sydney, New South Wales, Australia

Suzanne J. Hand

Affiliation School of Biological, Earth and Environmental Sciences, University of New South Wales, Sydney, New South Wales, Australia

Competing Interests

The authors have the following interests: this research was partly supported by funds from Xstrata Copper Community Partnership Program North Queensland and Outback at Isa which were administered through the Australian Research Councils Linkage Projects Scheme. There are no patents, products in development or marketed products to declare. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Analyzed the data: KHB ABC. Wrote the paper: KHB ABC MA SJH. Carried out anatomical comparisons: KHB ABC. Processed majority of fossil specimens: KHB. Wrote majority of manuscript: KHB.