About the Authors

Sara E. Hester

Affiliations Department of Veterinary and Biomedical Sciences, The Pennsylvania State University, University Park, Pennsylvania, United States of America, Graduate Program in Biochemistry, Microbiology, and Molecular Biology, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Minghsun Lui

Affiliation Department of Microbiology, Immunology, and Molecular Genetics, David Geffen School of Medicine at UCLA, University of California Los Angeles, Los Angeles, California, United States of America

Tracy Nicholson

Affiliation National Animal Disease Center, Agricultural Research Service, United States Department of Agriculture, Ames, Iowa, United State of America

Daryl Nowacki

Affiliations Department of Veterinary and Biomedical Sciences, The Pennsylvania State University, University Park, Pennsylvania, United States of America, Graduate Program in Biochemistry, Microbiology, and Molecular Biology, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Eric T. Harvill

harvill@psu.edu

Affiliation Department of Veterinary and Biomedical Sciences, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SEH ML TN ETH. Performed the experiments: SEH ML TN DN. Analyzed the data: SEH ML TN DN. Contributed reagents/materials/analysis tools: TN ETH. Wrote the paper: SEH TN ETH.