About the Authors

Andrew J. Love

Current address: Cell and Molecular Sciences, James Hutton Research Institute, Invergowrie, Dundee, United Kingdom

Affiliation Plant Science Research Theme, School of Life Sciences and Institute of Molecular, Cellular and Systems Biology, College of Medical, Veterinary and Life Sciences, University of Glasgow, Glasgow, Scotland, United Kingdom

Chiara Geri

Affiliations Plant Science Research Theme, School of Life Sciences and Institute of Molecular, Cellular and Systems Biology, College of Medical, Veterinary and Life Sciences, University of Glasgow, Glasgow, Scotland, United Kingdom, Istituto di Biologia e Biotechnologia Agraria, Consiglio Nazionale Delle Richerche, Pisa, Italy

Janet Laird

Affiliation Plant Science Research Theme, School of Life Sciences and Institute of Molecular, Cellular and Systems Biology, College of Medical, Veterinary and Life Sciences, University of Glasgow, Glasgow, Scotland, United Kingdom

Craig Carr

Affiliation Plant Science Research Theme, School of Life Sciences and Institute of Molecular, Cellular and Systems Biology, College of Medical, Veterinary and Life Sciences, University of Glasgow, Glasgow, Scotland, United Kingdom

Byung-Wook Yun

Current address: Division of Plant Biosciences, School of Applied Biosciences, College of Agriculture & Life Science, Kyungpook National University, Daegu, South Korea

Affiliation Institute of Molecular Plant Sciences, University of Edinburgh, King's Buildings, Edinburgh, United Kingdom

Gary J. Loake

Affiliation Institute of Molecular Plant Sciences, University of Edinburgh, King's Buildings, Edinburgh, United Kingdom

Yasuomi Tada

Current address: Life Science Research Center, Kagawa University, Kagawa, Japan

Affiliation Department of Biology, Duke University, Durham, North Carolina, United States of America

Ari Sadanandom

Current address: School of Biological and Biomedical Sciences, Durham University, Durham, United Kingdom

Affiliation Plant Science Research Theme, School of Life Sciences and Institute of Molecular, Cellular and Systems Biology, College of Medical, Veterinary and Life Sciences, University of Glasgow, Glasgow, Scotland, United Kingdom

Joel J. Milner

joel.milner@glasgow.ac.uk

Affiliation Plant Science Research Theme, School of Life Sciences and Institute of Molecular, Cellular and Systems Biology, College of Medical, Veterinary and Life Sciences, University of Glasgow, Glasgow, Scotland, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JJM AJL GJL AS CG. Performed the experiments: AJL CG JL CC BWY YT JJM. Analyzed the data: JJM AJL AS CG JL. Contributed reagents/materials/analysis tools: YT. Wrote the paper: JJM AJL AS.