About the Authors

Fumiaki Itoi

Affiliations Laboratory for Genomic Reprogramming, Center for Developmental Biology, RIKEN, Kobe, Japan, The Asada Institute for Reproductive Medicine, Asada Ladies Clinic, Kasugai, Japan

Mikiko Tokoro

Affiliations Laboratory for Genomic Reprogramming, Center for Developmental Biology, RIKEN, Kobe, Japan, The Asada Institute for Reproductive Medicine, Asada Ladies Clinic, Kasugai, Japan

Yukari Terashita

Affiliations Laboratory for Genomic Reprogramming, Center for Developmental Biology, RIKEN, Kobe, Japan, Laboratory of Animal Reproduction, Graduate School of Agricultural Science, Tohoku University, Sendai, Japan

Kazuo Yamagata

Affiliation Center for Genetic Analysis of Biological Responses, Research Institute for Microbial Diseases, Osaka University, Suita, Japan

Noritaka Fukunaga

Affiliation The Asada Institute for Reproductive Medicine, Asada Ladies Clinic, Kasugai, Japan

Yoshimasa Asada

Affiliation The Asada Institute for Reproductive Medicine, Asada Ladies Clinic, Kasugai, Japan

Teruhiko Wakayama

teru@cdb.riken.jp

Affiliations Laboratory for Genomic Reprogramming, Center for Developmental Biology, RIKEN, Kobe, Japan, Faculty of Life and Environmental Sciences, University of Yamanashi, Yamanashi, Japan, Kofu, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TW FI KY YA. Performed the experiments: FI MT YT. Analyzed the data: FI NF. Contributed reagents/materials/analysis tools: TW. Wrote the paper: TW FI.