About the Authors

Zurab R. Tsetskhladze

zut555@gmail.com (ZRT); kcheng76@gmail.com (KCC)

Affiliations Jake Gittlen Cancer Research Foundation, Penn State Hershey College of Medicine, Hershey, Pennsylvania, United States of America, Division of Experimental Pathology, Department of Pathology, Penn State Hershey College of Medicine, Hershey, Pennsylvania, United States of America

Victor A. Canfield

Affiliation Department of Pharmacology, Penn State Hershey College of Medicine, Hershey, Pennsylvania, United States of America

Khai C. Ang

Affiliations Jake Gittlen Cancer Research Foundation, Penn State Hershey College of Medicine, Hershey, Pennsylvania, United States of America, Division of Experimental Pathology, Department of Pathology, Penn State Hershey College of Medicine, Hershey, Pennsylvania, United States of America

Steven M. Wentzel

Affiliations Jake Gittlen Cancer Research Foundation, Penn State Hershey College of Medicine, Hershey, Pennsylvania, United States of America, Division of Experimental Pathology, Department of Pathology, Penn State Hershey College of Medicine, Hershey, Pennsylvania, United States of America

Katherine P. Reid

Affiliations Jake Gittlen Cancer Research Foundation, Penn State Hershey College of Medicine, Hershey, Pennsylvania, United States of America, Division of Experimental Pathology, Department of Pathology, Penn State Hershey College of Medicine, Hershey, Pennsylvania, United States of America

Arthur S. Berg

Affiliation Department of Public Health Sciences, Penn State Hershey College of Medicine, Hershey, Pennsylvania, United States of America

Stephen L. Johnson

Affiliation Department of Genetics, Washington University Medical School, St. Louis, Missouri, United States of America

Koichi Kawakami

Affiliation Division of Molecular and Developmental Biology, National Institute of Genetics, Department of Genetics, Graduate University for Advanced Studies (Sokendai), Mishima, Shizuoka, Japan

Keith C. Cheng

zut555@gmail.com (ZRT); kcheng76@gmail.com (KCC)

Affiliations Jake Gittlen Cancer Research Foundation, Penn State Hershey College of Medicine, Hershey, Pennsylvania, United States of America, Division of Experimental Pathology, Department of Pathology, Penn State Hershey College of Medicine, Hershey, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KCC VAC ZRT. Performed the experiments: ZRT KCA KPR SMW. Analyzed the data: KCC ZRT VAC KCA SMW ASB. Contributed reagents/materials/analysis tools: KCC SLJ KK. Wrote the paper: ZRT KCC KPR.