About the Authors

Jenny A. Hodgson

jenny.hodgson@york.ac.uk

Affiliation Department of Biology, University of York, York, United Kingdom

Chris D. Thomas

Affiliation Department of Biology, University of York, York, United Kingdom

Calvin Dytham

Affiliation Department of Biology, University of York, York, United Kingdom

Justin M. J. Travis

Affiliation Institute of Biological and Environmental Sciences, University of Aberdeen, Aberdeen, United Kingdom

Stephen J. Cornell

Affiliation School of Biology, University of Leeds, Leeds, United Kingdom

Competing Interests

Co-author Stephen Cornell is a PLOS ONE Editorial Board member. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: JAH CDT CD JMJT SJC. Performed the experiments: JAH. Analyzed the data: JAH. Contributed reagents/materials/analysis tools: SJC. Wrote the paper: JAH. Edited the manuscript: JAH CDT CD JMJT SJC.