About the Authors

Takeaki Ishihara

Affiliation Plant Protection Division, National Agricultural Research Center, National Agriculture and Food Research Organization, Tsukuba, Ibaraki, Japan

Ichiro Mitsuhara

Affiliation Division of Plant Sciences, National Institute of Agrobiological Sciences, Tsukuba, Ibaraki, Japan

Hideki Takahashi

Affiliation Department of Life Science, Graduate School of Agricultural Science, Tohoku University, Sendai, Miyagi, Japan

Kazuhiro Nakaho

nakaho@affrc.go.jp

Affiliation Plant Protection Division, National Agricultural Research Center, National Agriculture and Food Research Organization, Tsukuba, Ibaraki, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TI KN. Performed the experiments: TI KN. Analyzed the data: TI IM KN. Contributed reagents/materials/analysis tools: TI IM HT KN. Wrote the paper: TI KN.