About the Authors

Sowmya Parameswaran

Affiliation Department of Ophthalmology and Visual Sciences, University of Nebraska Medical Center, Omaha, Nebraska, United States of America

Sudha Balasubramanian

Affiliation Department of Ophthalmology and Visual Sciences, University of Nebraska Medical Center, Omaha, Nebraska, United States of America

Norbert Babai

Affiliation Department of Ophthalmology and Visual Sciences, University of Nebraska Medical Center, Omaha, Nebraska, United States of America

Carolina B. DelDebbio

Affiliation Department of Ophthalmology and Visual Sciences, University of Nebraska Medical Center, Omaha, Nebraska, United States of America

Donald W. Harms

Affiliation Department of Genetics Cell Biology and Anatomy, University of Nebraska Medical Center, Omaha, Nebraska, United States of America

Channabasavaiah B. Gurumurthy

Affiliation Department of Genetics Cell Biology and Anatomy, University of Nebraska Medical Center, Omaha, Nebraska, United States of America

Mahendra S. Rao

Affiliation Center for Regenerative Medicine,National Institutes of Health, Bethesda, Maryland, United States of America

John G. Sharp

Affiliation Department of Genetics Cell Biology and Anatomy, University of Nebraska Medical Center, Omaha, Nebraska, United States of America

Iqbal Ahmad

iahmad@unmc.edu

Affiliation Department of Ophthalmology and Visual Sciences, University of Nebraska Medical Center, Omaha, Nebraska, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: IA. Performed the experiments: SP SB NB CBDD DWH. Analyzed the data: SP SB CBG MSR JGS. Wrote the paper: SP IA.