About the Authors

Aya Miyazaki

Contributed equally to this work with: Aya Miyazaki, Kentaro Ishii

Affiliation Department of Life Sciences (Biology), Graduate School of Arts and Sciences, the University of Tokyo, Tokyo, Japan

Kentaro Ishii

Contributed equally to this work with: Aya Miyazaki, Kentaro Ishii

Affiliation Department of Life Sciences (Biology), Graduate School of Arts and Sciences, the University of Tokyo, Tokyo, Japan

Satoshi Yamashita

Affiliation Department of Life Sciences (Biology), Graduate School of Arts and Sciences, the University of Tokyo, Tokyo, Japan

Susumu Nejigane

Affiliation Department of Life Sciences (Biology), Graduate School of Arts and Sciences, the University of Tokyo, Tokyo, Japan

Shinya Matsukawa

Affiliation Department of Life Sciences (Biology), Graduate School of Arts and Sciences, the University of Tokyo, Tokyo, Japan

Yuzuru Ito

Affiliation Research Center for Stem Cell Engineering, National Institute of Advanced Industrial Science and Technology (AIST), Tsukuba City, Ibaraki, Japan

Yasuko Onuma

Affiliation Research Center for Stem Cell Engineering, National Institute of Advanced Industrial Science and Technology (AIST), Tsukuba City, Ibaraki, Japan

Makoto Asashima

Affiliations Department of Life Sciences (Biology), Graduate School of Arts and Sciences, the University of Tokyo, Tokyo, Japan, Research Center for Stem Cell Engineering, National Institute of Advanced Industrial Science and Technology (AIST), Tsukuba City, Ibaraki, Japan

Tatsuo Michiue

tmichiue@bio.c.u-tokyo.ac.jp

Affiliation Department of Life Sciences (Biology), Graduate School of Arts and Sciences, the University of Tokyo, Tokyo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TM. Performed the experiments: TM AM KI SY SN SM. Analyzed the data: TM AM YI YO SM. Contributed reagents/materials/analysis tools: TM YI YO MA SM. Wrote the paper: TM AM.