About the Authors

Hiroshi Nittono

nittono@hiroshima-u.ac.jp

Affiliation Graduate School of Integrated Arts and Sciences, Hiroshima University, Higashi-Hiroshima, Hiroshima, Japan

Michiko Fukushima

Affiliation Graduate School of Integrated Arts and Sciences, Hiroshima University, Higashi-Hiroshima, Hiroshima, Japan

Akihiro Yano

Affiliation Graduate School of Integrated Arts and Sciences, Hiroshima University, Higashi-Hiroshima, Hiroshima, Japan

Hiroki Moriya

Affiliation Graduate School of Integrated Arts and Sciences, Hiroshima University, Higashi-Hiroshima, Hiroshima, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HN MF AY HM. Performed the experiments: MF AY HM. Analyzed the data: HN MF AY HM. Wrote the paper: HN. Organized the research project: HN.