About the Authors

Richard H. Heineman

Affiliation Section of Integrative Biology, University of Texas at Austin, Austin, Texas, United States of America

Sam P. Brown

sam.brown@ed.ac.uk

Affiliations Section of Integrative Biology, University of Texas at Austin, Austin, Texas, United States of America, Centre for Infection, Immunity and Evolution, University of Edinburgh, Edinburgh, United Kingdom

Competing Interests

Sam Paul Brown is a PLOS ONE Editorial Board member. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: SPB RHH. Performed the experiments: SPB RHH. Analyzed the data: SPB RHH. Contributed reagents/materials/analysis tools: SPB RHH. Wrote the paper: SPB RHH.