About the Authors

Jesse W. Breinholt

jessebreinholt@gmail.com

Affiliation Department of Biology, Brigham Young University, Provo, Utah, United States of America

Megan L. Porter

Affiliation Department of Biology, University of South Dakota, Vermillion, South Dakota, United States of America

Keith A. Crandall

Affiliations Department of Biology, Brigham Young University, Provo, Utah, United States of America, Computational Biology Institute, George Washington University, Ashburn, Virginia, United States of America

Competing Interests

Co-author Keith A. Crandall is a PLOS ONE Editorial Board member. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: KAC JWB MLP. Performed the experiments: JWB MLP. Analyzed the data: JWB KAC. Contributed reagents/materials/analysis tools: KAC. Wrote the paper: JWB KAC MLP.