About the Authors

Sentaro Kusuhara

Contributed equally to this work with: Sentaro Kusuhara, Yoko Fukushima

Affiliations Laboratory for Stem Cell Biology, RIKEN Center for Developmental Biology, Kobe, Japan, Division of Ophthalmology, Department of Surgery, Kobe University Graduate School of Medicine, Kobe, Japan

Yoko Fukushima

Contributed equally to this work with: Sentaro Kusuhara, Yoko Fukushima

Affiliations Laboratory for Stem Cell Biology, RIKEN Center for Developmental Biology, Kobe, Japan, Division of Vascular Biology, Department of Physiology and Cell Biology, Kobe University Graduate School of Medicine, Kobe, Japan, Department of Ophthalmology, Osaka University Medical School, Osaka, Japan

Shigetomo Fukuhara

Affiliation Department of Cell Biology, National Cerebral and Cardiovascular Center Research Institute, Osaka, Japan

Lars Martin Jakt

Affiliation Laboratory for Stem Cell Biology, RIKEN Center for Developmental Biology, Kobe, Japan

Mitsuhiro Okada

Affiliation Laboratory for Stem Cell Biology, RIKEN Center for Developmental Biology, Kobe, Japan

Yuri Shimizu

Affiliation Laboratory for Stem Cell Biology, RIKEN Center for Developmental Biology, Kobe, Japan

Masayuki Hata

Affiliation Division of Vascular Biology, Department of Physiology and Cell Biology, Kobe University Graduate School of Medicine, Kobe, Japan

Kohji Nishida

Affiliation Department of Ophthalmology, Osaka University Medical School, Osaka, Japan

Akira Negi

Affiliation Division of Ophthalmology, Department of Surgery, Kobe University Graduate School of Medicine, Kobe, Japan

Masanori Hirashima

Affiliation Division of Vascular Biology, Department of Physiology and Cell Biology, Kobe University Graduate School of Medicine, Kobe, Japan

Naoki Mochizuki

Affiliation Department of Cell Biology, National Cerebral and Cardiovascular Center Research Institute, Osaka, Japan

Shin-Ichi Nishikawa

Affiliation Laboratory for Stem Cell Biology, RIKEN Center for Developmental Biology, Kobe, Japan

Akiyoshi Uemura

auemura@med.kobe-u.ac.jp

Affiliations Laboratory for Stem Cell Biology, RIKEN Center for Developmental Biology, Kobe, Japan, Division of Ophthalmology, Department of Surgery, Kobe University Graduate School of Medicine, Kobe, Japan, Division of Vascular Biology, Department of Physiology and Cell Biology, Kobe University Graduate School of Medicine, Kobe, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SK YF KN AN M. Hirashima SN AU. Performed the experiments: SK YF MO YS M. Hata AU. Analyzed the data: SK YF LMJ AU. Contributed reagents/materials/analysis tools: SF NM. Wrote the paper: AU.