About the Authors

Andrew T. McDonald

mcandr@sas.upenn.edu

Affiliation Department of Earth and Environmental Science, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

John Bird

Affiliation College of Eastern Utah Prehistoric Museum, Price, Utah, United States of America

James I. Kirkland

Affiliation Utah Geological Survey, Salt Lake City, Utah, United States of America

Peter Dodson

Affiliations Department of Earth and Environmental Science, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Competing Interests

Peter Dodson currently serves as an editor for PLOS ONE. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: ATM JB JIK. Performed the experiments: ATM JB JIK. Analyzed the data: ATM JB JIK. Contributed reagents/materials/analysis tools: ATM JB JIK PD. Wrote the paper: ATM.