About the Authors

Evan Maxwell

Affiliations Bioinformatics Program, Center for Biomedical Mass Spectrometry, Department, of Biochemistry, Boston University, Boston, Massachusetts, United States of America, Genome Technology Branch, National Human Genome Research Institute, National Institutes of Health, Bethesda, Maryland, United States of America

Yan Tan

Affiliations Bioinformatics Program, Center for Biomedical Mass Spectrometry, Department, of Biochemistry, Boston University, Boston, Massachusetts, United States of America, Broad Institute of Massachusetts Institute of Technology and Harvard, Cambridge, Massachusetts, United States of America

Yuxiang Tan

Affiliation Bioinformatics Program, Center for Biomedical Mass Spectrometry, Department, of Biochemistry, Boston University, Boston, Massachusetts, United States of America

Han Hu

Affiliation Bioinformatics Program, Center for Biomedical Mass Spectrometry, Department, of Biochemistry, Boston University, Boston, Massachusetts, United States of America

Gary Benson

Affiliation Bioinformatics Program, Center for Biomedical Mass Spectrometry, Department, of Biochemistry, Boston University, Boston, Massachusetts, United States of America

Konstantin Aizikov

Affiliation Center for Biomedical Mass Spectrometry, Department, of Biochemistry, Boston University, Boston, Massachusetts, United States of America

Shannon Conley

Affiliation Center for Biomedical Mass Spectrometry, Department, of Biochemistry, Boston University, Boston, Massachusetts, United States of America

Gregory O. Staples

Affiliation Center for Biomedical Mass Spectrometry, Department, of Biochemistry, Boston University, Boston, Massachusetts, United States of America

Gordon W. Slysz

Affiliation Biological Sciences Division and Environmental Molecular Sciences Laboratory, Pacific Northwest National Laboratory, Richland, Washington, United States of America

Richard D. Smith

Affiliation Biological Sciences Division and Environmental Molecular Sciences Laboratory, Pacific Northwest National Laboratory, Richland, Washington, United States of America

Joseph Zaia

jzaia@bu.edu

Affiliation Center for Biomedical Mass Spectrometry, Department, of Biochemistry, Boston University, Boston, Massachusetts, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JZ EM. Performed the experiments: YanT YuxiangT EM G. Staples. Analyzed the data: YanT YuxiangT EM JZ. Contributed reagents/materials/analysis tools: G. Staples RDS G. Slysz. Wrote the paper: JZ EM YanT YuxiangT. Developed early version of the software: KA SC. Assisted with the programming of GlycReSoft: HH. Guided the development of the bioinformatics algorithm: GB.