About the Authors

A. James Watkins

Contributed equally to this work with: A. James Watkins, Rifat A. Hamoudi

Affiliation Division of Molecular Histopathology, Department of Pathology, University of Cambridge, Cambridge, United Kingdom

Rifat A. Hamoudi

Contributed equally to this work with: A. James Watkins, Rifat A. Hamoudi

Affiliation Division of Molecular Histopathology, Department of Pathology, University of Cambridge, Cambridge, United Kingdom

Naiyan Zeng

Affiliation Division of Molecular Histopathology, Department of Pathology, University of Cambridge, Cambridge, United Kingdom

Qingguo Yan

Affiliations Division of Molecular Histopathology, Department of Pathology, University of Cambridge, Cambridge, United Kingdom, State Key Laboratory of Cancer Biology, Department of Pathology, Xijing Hospital, Xian, People’s Republic of China

Yuanxue Huang

Affiliation Division of Molecular Histopathology, Department of Pathology, University of Cambridge, Cambridge, United Kingdom

Hongxiang Liu

Affiliation Department of Histopathology, Addenbrooke’s Hospital, Cambridge, United Kingdom

Jianzhong Zhang

Affiliation Department of Pathology, 306 Hospital, Beijing, People’s Republic of China

Esteban Braggio

Affiliation Comprehensive Cancer Centre, Mayo Clinic in Arizona, Scottsdale, Arizona, United States of America

Rafael Fonseca

Affiliation Division of Hematology, Cancer Center – Oncology, Mayo Clinic in Arizona, Scottsdale, Arizona, United States of America

Laurence de Leval

Affiliation Institut Universitaire De Pathologie, Lausanne, Switzerland

Peter G. Isaacson

Affiliation Department of Histopathology, University College London Hospital, London, United Kingdom

Andrew Wotherspoon

Affiliation Department of Histopathology, Royal Marsden Hospital, London, United Kingdom

Ellen D. McPhail

Affiliation Department of Laboratory Medicine and Pathology, Mayo Clinic, Rochester, Minnesota, United States of America

Ahmet Dogan

These authors also contributed equally to this work.

Affiliation Department of Laboratory Medicine and Pathology, Mayo Clinic, Rochester, Minnesota, United States of America

Ming-Qing Du

mqd20@cam.ac.uk

These authors also contributed equally to this work.

Affiliations Division of Molecular Histopathology, Department of Pathology, University of Cambridge, Cambridge, United Kingdom, Department of Histopathology, Addenbrooke’s Hospital, Cambridge, United Kingdom

Competing Interests

All authors have declared that no competing interests exist. The Mayo Clinic is a nonprofit medical center and therefore should not be considered a commercial company. All the authors from Mayo Clinic have declared that no competing interests exist for this manuscript.

Author Contributions

Conceived and designed the experiments: AJW MQD. Performed the experiments: AJW RAH NZ QY YH EB RF EDM AD. Analyzed the data: AJW RAH EB RF EDM AD MQD. Contributed reagents/materials/analysis tools: HL JZ LdL PGI AW EB RF EDM AD. Wrote the paper: AJW MQD.