About the Authors

Lin Xu

Contributed equally to this work with: Lin Xu, Xia He, Ding-mei Zhang

Affiliations Department of Microbiology, Zhongshan School of Medicine, Sun Yat-sen University, Guangzhou, People's Republic of China, Key Laboratory of Tropical Disease Control, Ministry of Education, Sun Yat-Sen University, Guangzhou, People's Republic of China, Sun Yat-sen University – University of Hong Kong Joint Laboratory of Infectious Disease Surveillance, Sun Yat-sen University, Guangzhou, People's Republic of China

Xia He

Contributed equally to this work with: Lin Xu, Xia He, Ding-mei Zhang

Affiliations Department of Microbiology, Zhongshan School of Medicine, Sun Yat-sen University, Guangzhou, People's Republic of China, Key Laboratory of Tropical Disease Control, Ministry of Education, Sun Yat-Sen University, Guangzhou, People's Republic of China, Sun Yat-sen University – University of Hong Kong Joint Laboratory of Infectious Disease Surveillance, Sun Yat-sen University, Guangzhou, People's Republic of China

Ding-mei Zhang

Contributed equally to this work with: Lin Xu, Xia He, Ding-mei Zhang

Affiliations Department of Microbiology, Zhongshan School of Medicine, Sun Yat-sen University, Guangzhou, People's Republic of China, Key Laboratory of Tropical Disease Control, Ministry of Education, Sun Yat-Sen University, Guangzhou, People's Republic of China, Sun Yat-sen University – University of Hong Kong Joint Laboratory of Infectious Disease Surveillance, Sun Yat-sen University, Guangzhou, People's Republic of China

Fa-shen Feng

Affiliations Department of Microbiology, Zhongshan School of Medicine, Sun Yat-sen University, Guangzhou, People's Republic of China, Key Laboratory of Tropical Disease Control, Ministry of Education, Sun Yat-Sen University, Guangzhou, People's Republic of China, Sun Yat-sen University – University of Hong Kong Joint Laboratory of Infectious Disease Surveillance, Sun Yat-sen University, Guangzhou, People's Republic of China

Zhu Wang

Affiliations Department of Microbiology, Zhongshan School of Medicine, Sun Yat-sen University, Guangzhou, People's Republic of China, Key Laboratory of Tropical Disease Control, Ministry of Education, Sun Yat-Sen University, Guangzhou, People's Republic of China, Sun Yat-sen University – University of Hong Kong Joint Laboratory of Infectious Disease Surveillance, Sun Yat-sen University, Guangzhou, People's Republic of China

Lin-lin Guan

Affiliations Department of Microbiology, Zhongshan School of Medicine, Sun Yat-sen University, Guangzhou, People's Republic of China, Key Laboratory of Tropical Disease Control, Ministry of Education, Sun Yat-Sen University, Guangzhou, People's Republic of China, Sun Yat-sen University – University of Hong Kong Joint Laboratory of Infectious Disease Surveillance, Sun Yat-sen University, Guangzhou, People's Republic of China

Jue-heng Wu

Affiliations Department of Microbiology, Zhongshan School of Medicine, Sun Yat-sen University, Guangzhou, People's Republic of China, Key Laboratory of Tropical Disease Control, Ministry of Education, Sun Yat-Sen University, Guangzhou, People's Republic of China, Sun Yat-sen University – University of Hong Kong Joint Laboratory of Infectious Disease Surveillance, Sun Yat-sen University, Guangzhou, People's Republic of China

Rong Zhou

Affiliation State Key Laboratory of Respiratory Diseases, Guangzhou Medical University, Guangzhou, People's Republic of China

Bo-jian Zheng

Affiliations Sun Yat-sen University – University of Hong Kong Joint Laboratory of Infectious Disease Surveillance, Sun Yat-sen University, Guangzhou, People's Republic of China, Department of Microbiology, University of Hong Kong, Hong Kong SAR, China

Kwok-yung Yuen

Affiliations Sun Yat-sen University – University of Hong Kong Joint Laboratory of Infectious Disease Surveillance, Sun Yat-sen University, Guangzhou, People's Republic of China, Department of Microbiology, University of Hong Kong, Hong Kong SAR, China

Meng-feng Li

Affiliations Department of Microbiology, Zhongshan School of Medicine, Sun Yat-sen University, Guangzhou, People's Republic of China, Key Laboratory of Tropical Disease Control, Ministry of Education, Sun Yat-Sen University, Guangzhou, People's Republic of China, Sun Yat-sen University – University of Hong Kong Joint Laboratory of Infectious Disease Surveillance, Sun Yat-sen University, Guangzhou, People's Republic of China

Kai-yuan Cao

caoky@mail.sysu.edu.cn

Affiliations Department of Microbiology, Zhongshan School of Medicine, Sun Yat-sen University, Guangzhou, People's Republic of China, Key Laboratory of Tropical Disease Control, Ministry of Education, Sun Yat-Sen University, Guangzhou, People's Republic of China, Sun Yat-sen University – University of Hong Kong Joint Laboratory of Infectious Disease Surveillance, Sun Yat-sen University, Guangzhou, People's Republic of China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: K-yC LX. Performed the experiments: LX XH D-mZ F-sF ZW L-lG. Analyzed the data: LX XH K-yY M-fL. Contributed reagents/materials/analysis tools: RZ B-jZ J-hW M-fL. Wrote the paper: LX XH M-fL K-yY.