About the Authors

Clémence Lesimple

clemence.lesimple@univ-rennes1.fr

Affiliation Université de Rennes 1, Laboratoire d’éthologie Animale et Humaine EthoS - UMR CNRS 6552, Station biologique, Paimpont, France

Carole Fureix

Current address: Animal Behaviour and Welfare Research Group, Animal and Poultry Science Department, University of Guelph, Guelph, Ontario, Canada

Affiliation Université de Rennes 1, Laboratoire d’éthologie Animale et Humaine - UMR CNRS 6552, Campus de Beaulieu, Rennes, France

Emmanuel De Margerie

Affiliation Université de Rennes 1, Laboratoire d’éthologie Animale et Humaine - UMR CNRS 6552, Campus de Beaulieu, Rennes, France

Emilie Sénèque

Affiliation Université de Rennes 1, Laboratoire d’éthologie Animale et Humaine - UMR CNRS 6552, Campus de Beaulieu, Rennes, France

Hervé Menguy

Affiliation Chiropractic Practice, St Jacques de la Lande, France

Martine Hausberger

Affiliations Université de Rennes 1, Laboratoire d’éthologie Animale et Humaine EthoS - UMR CNRS 6552, Station biologique, Paimpont, France, Université de Rennes 1, Laboratoire d’éthologie Animale et Humaine - UMR CNRS 6552, Campus de Beaulieu, Rennes, France

Competing Interests

MH is an Editor for PLoS ONE. This does not alter the authors’ adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: MH CL CF. Performed the experiments: CL CF HM. Analyzed the data: CL MH. Contributed reagents/materials/analysis tools: EDM ES. Wrote the paper: CL MH.