About the Authors

Helen Downie

Affiliations The James Hutton Institute, Invergowrie, Dundee, United Kingdom, The SIMBIOS Centre, University of Abertay Dundee, Bell Street, Dundee, United Kingdom

Nicola Holden

Affiliation The James Hutton Institute, Invergowrie, Dundee, United Kingdom

Wilfred Otten

Affiliation The SIMBIOS Centre, University of Abertay Dundee, Bell Street, Dundee, United Kingdom

Andrew J. Spiers

Affiliation The SIMBIOS Centre, University of Abertay Dundee, Bell Street, Dundee, United Kingdom

Tracy A. Valentine

Affiliation The James Hutton Institute, Invergowrie, Dundee, United Kingdom

Lionel X. Dupuy

Lionel.Dupuy@Hutton.ac.uk

Affiliation The James Hutton Institute, Invergowrie, Dundee, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HD NH WO AJS TAV LXD. Performed the experiments: HD. Analyzed the data: HD LXD. Contributed reagents/materials/analysis tools: HD LXD. Wrote the paper: HD NH WO AJS TAV LXD.