About the Authors

Keigo Inami

Affiliation Graduate School of Agriculture, Tokyo University of Agriculture and Technology (TUAT), Fuchu, Japan

Chizu Yoshioka-Akiyama

Affiliation Graduate School of Agriculture, Tokyo University of Agriculture and Technology (TUAT), Fuchu, Japan

Yasuaki Morita

Affiliation Kochi Agricultural Research Center, Nangoku, Japan

Mutsuko Yamasaki

Affiliations Graduate School of Agriculture, Tokyo University of Agriculture and Technology (TUAT), Fuchu, Japan, Kochi Agricultural Research Center, Nangoku, Japan

Tohru Teraoka

Affiliation Graduate School of Agriculture, Tokyo University of Agriculture and Technology (TUAT), Fuchu, Japan

Tsutomu Arie

arie@cc.tuat.ac.jp

Affiliation Graduate School of Agriculture, Tokyo University of Agriculture and Technology (TUAT), Fuchu, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KI TA. Performed the experiments: KI CYA TA. Analyzed the data: KI TA. Contributed reagents/materials/analysis tools: YM MY. Wrote the paper: KI TT TA.