About the Authors

Joel W. Graff

Affiliations Department of Medicine, University of Iowa, Iowa City, Iowa, United States of America, Iowa City Veteran Affairs Medical Center, Iowa City, Iowa, United States of America

Linda S. Powers

Affiliation Department of Medicine, University of Iowa, Iowa City, Iowa, United States of America

Anne M. Dickson

Current address: Life Technologies, Eugene, Oregon, United States of America

Affiliation Department of Medicine, University of Iowa, Iowa City, Iowa, United States of America

Jongkwang Kim

Affiliation Department of Medicine, University of Iowa, Iowa City, Iowa, United States of America

Anna C. Reisetter

Affiliation Department of Medicine, University of Iowa, Iowa City, Iowa, United States of America

Ihab H. Hassan

Affiliation Department of Medicine, University of Iowa, Iowa City, Iowa, United States of America

Karol Kremens

Affiliation Department of Medicine, University of Iowa, Iowa City, Iowa, United States of America

Thomas J. Gross

Affiliation Department of Medicine, University of Iowa, Iowa City, Iowa, United States of America

Mary E. Wilson

Contributed equally to this work with: Mary E. Wilson, Martha M. Monick

Affiliations Department of Medicine, University of Iowa, Iowa City, Iowa, United States of America, Iowa City Veteran Affairs Medical Center, Iowa City, Iowa, United States of America, Department of Microbiology, University of Iowa, Iowa City, Iowa, United States of America

Martha M. Monick

Contributed equally to this work with: Mary E. Wilson, Martha M. Monick

martha-monick@uiowa.edu

Affiliation Department of Medicine, University of Iowa, Iowa City, Iowa, United States of America

Competing Interests

The authors have read the journal’s policy and have the following conflicts. Author A.D. is now employed at Life Technologies. This does not alter the authors’ adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: JG TG MW MM. Performed the experiments: LP AD AR IH KK. Analyzed the data: JG AD JK MM. Contributed reagents/materials/analysis tools: JK. Wrote the paper: JG MW MM.