About the Authors

Justin M. Greene

Affiliation Department of Pathology and Laboratory Medicine, University of Wisconsin-Madison, Madison, Wisconsin, United States of America

Emily N. Chin

Affiliation Department of Cellular and Molecular Biology, University of Wisconsin-Madison, Madison, Wisconsin, United States of America

Melisa L. Budde

Affiliation Wisconsin National Primate Research Center, University of Wisconsin-Madison, Madison, Wisconsin, United States of America

Jennifer J. Lhost

Affiliation Department of Pathology and Laboratory Medicine, University of Wisconsin-Madison, Madison, Wisconsin, United States of America

Paul J. Hines

Affiliation Wisconsin National Primate Research Center, University of Wisconsin-Madison, Madison, Wisconsin, United States of America

Benjamin J. Burwitz

Affiliation Wisconsin National Primate Research Center, University of Wisconsin-Madison, Madison, Wisconsin, United States of America

Karl W. Broman

Affiliation Department of Biostatistics and Medical Informatics, University of Wisconsin-Madison, Madison, Wisconsin, United States of America

Jennifer E. Nelson

Affiliation Wisconsin National Primate Research Center, University of Wisconsin-Madison, Madison, Wisconsin, United States of America

Thomas C. Friedrich

Affiliations Wisconsin National Primate Research Center, University of Wisconsin-Madison, Madison, Wisconsin, United States of America, Department of Pathobiological Sciences, University of Wisconsin, Madison, Wisconsin, United States of America

David H. O’Connor

david.h.oconnor@gmail.com

Affiliations Department of Pathology and Laboratory Medicine, University of Wisconsin-Madison, Madison, Wisconsin, United States of America, Department of Cellular and Molecular Biology, University of Wisconsin-Madison, Madison, Wisconsin, United States of America, Wisconsin National Primate Research Center, University of Wisconsin-Madison, Madison, Wisconsin, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: DHO JMG JJL ENC. Performed the experiments: ENC MLB JJL PJH BJB JEN. Analyzed the data: JMG KWB TCF. Contributed reagents/materials/analysis tools: TCF KWB. Wrote the paper: JMG DHO TCF.