About the Authors

Courtney S. Onodera

Current address: Institute for Human Genetics, University of California San Francisco, San Francisco, California, United States of America

Affiliation Department of Biomolecular Engineering, University of California Santa Cruz, Santa Cruz, California, United States of America

Jason G. Underwood

Current address: Pacific Biosciences, Menlo Park, California, United States of America

Affiliations Department of Biomolecular Engineering, University of California Santa Cruz, Santa Cruz, California, United States of America, Center for Biomolecular Sciences and Engineering, University of California Santa Cruz, Santa Cruz, California, United States of America, Howard Hughes Medical Institute, University of California Santa Cruz, Santa Cruz, California, United States of America

Sol Katzman

Affiliations Department of Biomolecular Engineering, University of California Santa Cruz, Santa Cruz, California, United States of America, Center for Biomolecular Sciences and Engineering, University of California Santa Cruz, Santa Cruz, California, United States of America

Frank Jacobs

Affiliations Department of Biomolecular Engineering, University of California Santa Cruz, Santa Cruz, California, United States of America, Center for Biomolecular Sciences and Engineering, University of California Santa Cruz, Santa Cruz, California, United States of America

David Greenberg

Affiliations Department of Biomolecular Engineering, University of California Santa Cruz, Santa Cruz, California, United States of America, Department of Molecular, Cell and Developmental Biology, University of California Santa Cruz, Santa Cruz, California, United States of America

Sofie R. Salama

ssalama@soe.ucsc.edu

Affiliations Department of Biomolecular Engineering, University of California Santa Cruz, Santa Cruz, California, United States of America, Center for Biomolecular Sciences and Engineering, University of California Santa Cruz, Santa Cruz, California, United States of America, Howard Hughes Medical Institute, University of California Santa Cruz, Santa Cruz, California, United States of America

David Haussler

Affiliations Department of Biomolecular Engineering, University of California Santa Cruz, Santa Cruz, California, United States of America, Center for Biomolecular Sciences and Engineering, University of California Santa Cruz, Santa Cruz, California, United States of America, Howard Hughes Medical Institute, University of California Santa Cruz, Santa Cruz, California, United States of America

Competing Interests

JGU is currently employed at Pacific Biosciences. This does not alter the authors' adherance to all the PLoS ONe policies on sharing data and materials. All the other authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CSO SRS DH. Performed the experiments: CSO JGU FJ DBG. Analyzed the data: CSO SK. Contributed reagents/materials/analysis tools: FJ. Wrote the paper: CSO SRS.