About the Authors

Camille Szmaragd

camille.szmaragd@bristol.ac.uk

Affiliation School of Veterinary Sciences, University of Bristol, Langford, Devon, United Kingdom

Laura E. Green

Affiliation School of Life Sciences, University of Warwick, Coventry, West Midlands, United Kingdom

Graham F. Medley

Affiliation School of Life Sciences, University of Warwick, Coventry, West Midlands, United Kingdom

William J. Browne

Affiliation School of Veterinary Sciences, University of Bristol, Langford, Devon, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CS LEG GFM WJB. Performed the experiments: CS. Analyzed the data: CS. Contributed reagents/materials/analysis tools: CS LEG GFM WJB. Wrote the paper: CS LEG GFM WJB. Designed and coded the model algorithm: CS WJB.