About the Authors

Guey Shuang Wu

Affiliation Department of Ophthalmology, Doheny Eye Institute, Keck School of Medicine, University of Southern California, Los Angeles, California

Meisheng Jiang

Affiliation Department of Molecular and Medical Pharmacology, David Geffen School of Medicine, University of California Los Angeles, Los Angeles, California

Yi-Hsin Liu

Affiliation Department of Ophthalmology, Doheny Eye Institute, Keck School of Medicine, University of Southern California, Los Angeles, California

Yoshiko Nagaoka

Affiliation Department of Molecular and Medical Pharmacology, David Geffen School of Medicine, University of California Los Angeles, Los Angeles, California

Narsing A. Rao

nrao@usc.edu

Affiliation Department of Ophthalmology, Doheny Eye Institute, Keck School of Medicine, University of Southern California, Los Angeles, California

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: GSW MJ YHL YN NAR. Performed the experiments: GSW MJ YHL YN. Analyzed the data: GSW NAR. Contributed reagents/materials/analysis tools: GSW MJ NAR. Wrote the paper: GSW.