About the Authors

Chris R. Smith

crsmith.ant@gmail.com

Affiliations Department of Biology, Earlham College, Richmond, Indiana, United States of America, Center for Social Dynamics and Complexity, School of Life Sciences, Arizona State University, Tempe, Arizona, United States of America

Navdeep S. Mutti

Affiliation Department of Biology, Earlham College, Richmond, Indiana, United States of America

W. Cameron Jasper

Affiliation Department of Biology, San Francisco State University, San Francisco, California, United States of America

Agni Naidu

Affiliation Department of Biology, San Francisco State University, San Francisco, California, United States of America

Christopher D. Smith

Affiliation Department of Biology, San Francisco State University, San Francisco, California, United States of America

Jürgen Gadau

Affiliation Department of Biology, Earlham College, Richmond, Indiana, United States of America

Competing Interests

The research that is presented in this article was performed while N. S. Mutti was at the Arizona State University, Tempe, AZ. N. S. Mutti is currently employed at DuPont, therefore Dr. Mutti's current contact information was updated to reflect change of employment. DuPont played no role in designing, conducting or funding the research presented in the article, therefore it won't alter the authors’ adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: CRS NSM CDS JG. Performed the experiments: CRS NSM WCJ AN CDS. Analyzed the data: CRS NSM WCJ AN CDS JG. Contributed reagents/materials/analysis tools: CRS NSM CDS JG. Wrote the paper: CRS NSM CDS WCJ JG.