About the Authors

Masaya Fukushi

mfukushi@hiroshima-u.ac.jp

Affiliations Disease Control and Prevention Center, National Center for Global Health and Medicine, Tokyo, Japan, Deputy Director-General's Laboratory, Research Institute, National Center for Global Health and Medicine, Tokyo, Japan, Department of Virology, Institute of Biomedical and Health Sciences, Hiroshima University, Hiroshima, Japan

Makoto Yamashita

Affiliation Biological Research Laboratories, Daiichi Sankyo Co., Ltd., Tokyo, Japan

Tohru Miyoshi-Akiyama

Affiliation Department of Infectious Diseases, Research Institute, National Center for Global Health and Medicine, Tokyo, Japan

Shuku Kubo

Affiliation Biological Research Laboratories, Daiichi Sankyo Co., Ltd., Tokyo, Japan

Kenji Yamamoto

Affiliation Deputy Director-General's Laboratory, Research Institute, National Center for Global Health and Medicine, Tokyo, Japan

Koichiro Kudo

Affiliation Disease Control and Prevention Center, National Center for Global Health and Medicine, Tokyo, Japan

Competing Interests

The authors declare that Makoto Yamashita and Shuku Kubo are employed by Biological Research Laboratories, Daiichi Sankyo Co., Ltd. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: MF MY KK. Performed the experiments: MF MY TMA SK. Analyzed the data: MF MY. Contributed reagents/materials/analysis tools: KY. Wrote the paper: MF MY KK.