About the Authors

Lyubov Popova

Affiliations Department of Biochemistry & Molecular Biology, University of Oklahoma Health Sciences Center, Oklahoma City, Oklahoma, United States of America, Department of Chemistry and Biochemistry, University of Oklahoma, Norman, Oklahoma, United States of America

Kenneth Smith

Affiliation Arthritis and Clinical Immunology Research Program, Oklahoma Medical Research Foundation, Oklahoma City, Oklahoma, United States of America

Ann H. West

Affiliation Department of Chemistry and Biochemistry, University of Oklahoma, Norman, Oklahoma, United States of America

Patrick C. Wilson

Affiliations Arthritis and Clinical Immunology Research Program, Oklahoma Medical Research Foundation, Oklahoma City, Oklahoma, United States of America, Immunobiology and Cancer Program, Oklahoma Medical Research Foundation, Oklahoma City, Oklahoma, United States of America, Section of Rheumatology, Department of Medicine and Committee on Immunology, University of Chicago, Chicago, Illinois, United States of America

Judith A. James

Affiliation Arthritis and Clinical Immunology Research Program, Oklahoma Medical Research Foundation, Oklahoma City, Oklahoma, United States of America

Linda F. Thompson

Affiliation Immunobiology and Cancer Program, Oklahoma Medical Research Foundation, Oklahoma City, Oklahoma, United States of America

Gillian M. Air

gillian-air@ouhsc.edu

Affiliation Department of Biochemistry & Molecular Biology, University of Oklahoma Health Sciences Center, Oklahoma City, Oklahoma, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: LP GMA. Performed the experiments: LP. Analyzed the data: LP AHW GMA. Contributed reagents/materials/analysis tools: LFT JAJ KS PCW. Wrote the paper: LP GMA. Obtained plasma samples: LFT JAJ. Characterized and purified monoclonal antibodies: PCW KS.