About the Authors

Dena G. Hernandez

Affiliation Laboratory of Neurogenetics, National Institute on Aging, National Institutes of Health, Bethesda, Maryland, United States of America

Michael A. Nalls

Affiliation Laboratory of Neurogenetics, National Institute on Aging, National Institutes of Health, Bethesda, Maryland, United States of America

Pauli Ylikotila

Affiliation Department of Neurology, Turku University Hospital, Turku, Finland

Margaux Keller

Affiliation Laboratory of Neurogenetics, National Institute on Aging, National Institutes of Health, Bethesda, Maryland, United States of America

John A. Hardy

Affiliation Reta Lilla Weston Laboratories and Departments of Molecular Neuroscience, UCL Institute of Neurology, Queen Square, London, United Kingdom

Kari Majamaa

Affiliation Institute of Clinical Medicine, Department of Neurology, University of Oulu, Oulu, Finland

Andrew B. Singleton

singleta@mail.nih.gov

Affiliation Laboratory of Neurogenetics, National Institute on Aging, National Institutes of Health, Bethesda, Maryland, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JH MN AS. Performed the experiments: DH. Analyzed the data: MN MK. Contributed reagents/materials/analysis tools: PY KM. Wrote the paper: AS DH KM MN PY MK.