About the Authors

Philip Francis Thomsen

Contributed equally to this work with: Philip Francis Thomsen, Jos Kielgast

pfthomsen@snm.ku.dk (PFT); ewillerslev@snm.ku.dk (EW)

Affiliation Centre for GeoGenetics, Natural History Museum of Denmark, University of Copenhagen, Øster Voldgade, Copenhagen, Denmark

Jos Kielgast

Contributed equally to this work with: Philip Francis Thomsen, Jos Kielgast

Affiliation Centre for GeoGenetics, Natural History Museum of Denmark, University of Copenhagen, Øster Voldgade, Copenhagen, Denmark

Lars Lønsmann Iversen

Affiliation Freshwater Biology Section, Department of Biology, University of Copenhagen, Helsingørgade, Hillerød, Denmark

Peter Rask Møller

Affiliation Vertebrate Department, Natural History Museum of Denmark, University of Copenhagen, Universitetsparken, Copenhagen, Denmark

Morten Rasmussen

Affiliation Centre for GeoGenetics, Natural History Museum of Denmark, University of Copenhagen, Øster Voldgade, Copenhagen, Denmark

Eske Willerslev

pfthomsen@snm.ku.dk (PFT); ewillerslev@snm.ku.dk (EW)

Affiliation Centre for GeoGenetics, Natural History Museum of Denmark, University of Copenhagen, Øster Voldgade, Copenhagen, Denmark

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: PFT JK EW. Performed the experiments: PFT JK PRM MR. Analyzed the data: PFT JK LLI PRM MR. Contributed reagents/materials/analysis tools: PFT JK LLI PRM MR. Wrote the paper: PFT JK PRM EW.