About the Authors

Katriina Tiira

Affiliations Research Programs Unit, Molecular Medicine, Department of Veterinary Biosciences, University of Helsinki, Helsinki, Finland, The Folkhälsan Research Center, Helsinki, Finland

Osmo Hakosalo

Affiliations Research Programs Unit, Molecular Medicine, Department of Veterinary Biosciences, University of Helsinki, Helsinki, Finland, The Folkhälsan Research Center, Helsinki, Finland

Lauri Kareinen

Affiliations Research Programs Unit, Molecular Medicine, Department of Veterinary Biosciences, University of Helsinki, Helsinki, Finland, The Folkhälsan Research Center, Helsinki, Finland

Anne Thomas

Affiliation Antagene, Animal Genetics Laboratory, Lyon, France

Anna Hielm-Björkman

Affiliation Department of Equine and Small Animal Medicine, University of Helsinki, Helsinki, Finland

Catherine Escriou

Affiliation National Veterinary School of Lyon, Lyon, France

Paul Arnold

Affiliation Program in Genetics and Genomic Biology, Hospital for Sick Children, Toronto, Ontario, Canada

Hannes Lohi

hannes.lohi@helsinki.fi

Affiliations Research Programs Unit, Molecular Medicine, Department of Veterinary Biosciences, University of Helsinki, Helsinki, Finland, The Folkhälsan Research Center, Helsinki, Finland

Competing Interests

AT works in a private Antagene company. This does not alter the authors’ adherence to all the PLoS ONE policies on sharing data and materials. The other authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KT HL. Performed the experiments: KT OH LK AT. Analyzed the data: KT OH AH-B. Contributed reagents/materials/analysis tools: HL AT PA CE. Wrote the paper: KT HL.