About the Authors

Kwonil Jung

Affiliation Food Animal Health Research Program, Ohio Agricultural Research and Development Center, Department of Veterinary Preventive Medicine, The Ohio State University, Wooster, Ohio, United States of America

Qiuhong Wang

Affiliation Food Animal Health Research Program, Ohio Agricultural Research and Development Center, Department of Veterinary Preventive Medicine, The Ohio State University, Wooster, Ohio, United States of America

Yunjeong Kim

Affiliation Diagnostic Medicine and Pathobiology, College of Veterinary Medicine, Kansas State University, Manhattan, Kansas, United States of America

Kelly Scheuer

Affiliation Food Animal Health Research Program, Ohio Agricultural Research and Development Center, Department of Veterinary Preventive Medicine, The Ohio State University, Wooster, Ohio, United States of America

Zhenwen Zhang

Affiliation Food Animal Health Research Program, Ohio Agricultural Research and Development Center, Department of Veterinary Preventive Medicine, The Ohio State University, Wooster, Ohio, United States of America

Quan Shen

Affiliation Food Animal Health Research Program, Ohio Agricultural Research and Development Center, Department of Veterinary Preventive Medicine, The Ohio State University, Wooster, Ohio, United States of America

Kyeong-Ok Chang

Affiliation Diagnostic Medicine and Pathobiology, College of Veterinary Medicine, Kansas State University, Manhattan, Kansas, United States of America

Linda J. Saif

saif.2@osu.edu

Affiliation Food Animal Health Research Program, Ohio Agricultural Research and Development Center, Department of Veterinary Preventive Medicine, The Ohio State University, Wooster, Ohio, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KJ QW YJK KOC LJS. Performed the experiments: KJ QW YJK KS ZZ QS. Analyzed the data: KJ QW LJS KOC. Contributed reagents/materials/analysis tools: KJ QW YJK KS ZZ QS LJS KOC. Wrote the paper: KJ QW KOC LJS.