About the Authors

Amy A. Briggs

Contributed equally to this work with: Amy A. Briggs, Hillary S. Young

Affiliation Department of Biology, Stanford University, Stanford, California, United States of America

Hillary S. Young

Contributed equally to this work with: Amy A. Briggs, Hillary S. Young

hyoung@fas.harvard.edu

Current address: Center for the Environment, Harvard University, Cambridge, Massachusetts, United States of America

Affiliation Department of Biology, Stanford University, Stanford, California, United States of America

Douglas J. McCauley

Current address: University of California, Berkeley, Department of Environmental Science, Policy, and Management, Berkeley, California, United States of America

Affiliation Department of Biology, Stanford University, Stanford, California, United States of America

Stacie A. Hathaway

Affiliation Western Ecological Research Center, U.S. Geological Survey, San Diego Field Station, San Diego, California, United States of America

Rodolfo Dirzo

Affiliation Department of Biology, Stanford University, Stanford, California, United States of America

Robert N. Fisher

Affiliation Western Ecological Research Center, U.S. Geological Survey, San Diego Field Station, San Diego, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HSY DJM RD. Performed the experiments: HSY AAB DJM SAH RNF. Analyzed the data: AAB HSY. Contributed reagents/materials/analysis tools: SAH RNF. Wrote the paper: AAB HSY SAH RNF DJM RD.