About the Authors

Zhiqiu Hu

Affiliation Department of Botany and Plant Sciences, University of California Reverside, Reverside, California, United States of America

Zhiquan Wang

Affiliation Department of Agricultural Food and Nutritional Science, University of Alberta, Edmonton, Alberta, Canada

Shizhong Xu

Shizhong.Xu@ucr.edu

Affiliation Department of Botany and Plant Sciences, University of California Reverside, Reverside, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SX. Performed the experiments: ZW. Analyzed the data: ZH. Contributed reagents/materials/analysis tools: ZW SX ZH. Wrote the paper: SX.