About the Authors

Audrey Maille

audrey.maille@univ-rennes1.fr

Affiliation UMR 6552 Ethos «Ethologie Animale et Humaine», University of Rennes 1 - CNRS, Station biologique, Paimpont, France

Lucie Engelhart

Affiliation UMR 6552 Ethos «Ethologie Animale et Humaine», University of Rennes 1 - CNRS, Station biologique, Paimpont, France

Marie Bourjade

Affiliation Research Center in the Psychology of Cognition, Language and Emotion, Aix-Marseille University, Aix-en-Provence, France

Catherine Blois-Heulin

Affiliation UMR 6552 Ethos «Ethologie Animale et Humaine», University of Rennes 1 - CNRS, Station biologique, Paimpont, France

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: AM LE CBH. Performed the experiments: LE. Analyzed the data: LE AM. Contributed reagents/materials/analysis tools: AM MB CBH. Wrote the paper: AM MB CBH.