About the Authors

Cornelia Schrauf

connyschrauf@chello.at

Affiliation Department of Behavioural Biology, University of Vienna, Vienna, Austria

Josep Call

Affiliation Department for Developmental and Comparative Psychology, Max Planck Institute for Evolutionary Anthropology, Leipzig, Germany

Koki Fuwa

Affiliation Great Ape Research Institute, Hayashibara Biochemical Laboratories, Tamano, Okayama, Japan

Satoshi Hirata

Affiliation Great Ape Research Institute, Hayashibara Biochemical Laboratories, Tamano, Okayama, Japan

Competing Interests

Two authors are employed by Hayashibara Biochemical Laboratories, Inc. However, the company had no role in study design, consultancy, data collection and analysis, decision to publish, or preperation of the manuscript. The authors declare their full adherence to all PLOS One policies on data sharing and materials.

Author Contributions

Conceived and designed the experiments: CS JC SH. Performed the experiments: KF SH. Analyzed the data: CS JC. Contributed reagents/materials/analysis tools: JC SH. Wrote the paper: CS JC.