About the Authors

Azadeh Shahsavar

Affiliation Department of Drug Design and Pharmacology, Faculty of Health and Medical Sciences, University of Copenhagen, Copenhagen, Denmark

Jette S. Kastrup

Affiliation Department of Drug Design and Pharmacology, Faculty of Health and Medical Sciences, University of Copenhagen, Copenhagen, Denmark

Elsebet Ø. Nielsen

Affiliation NeuroSearch A/S, Ballerup, Denmark

Jesper L. Kristensen

Affiliation Department of Drug Design and Pharmacology, Faculty of Health and Medical Sciences, University of Copenhagen, Copenhagen, Denmark

Michael Gajhede

Affiliation Department of Drug Design and Pharmacology, Faculty of Health and Medical Sciences, University of Copenhagen, Copenhagen, Denmark

Thomas Balle

thomas.balle@sydney.edu.au

Current address: Faculty of Pharmacy, The University of Sydney, Sydney, NSW, Australia

Affiliation Department of Drug Design and Pharmacology, Faculty of Health and Medical Sciences, University of Copenhagen, Copenhagen, Denmark

Competing Interests

EØN is employed by the company NeuroSearch A/S. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: AS JSK JLK MG TB. Performed the experiments: AS EØN. Analyzed the data: AS JSK EØN MG TB. Contributed reagents/materials/analysis tools: JLK. Wrote the paper: AS JSK TB.